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The African king of beasts has declined to < 35,000 persons busying 25 % of its historic scope. The state of affairs is most critical for the geographically stray populations in West Africa, where the species is considered regionally endangered. Promoting their preservation significance, recent molecular surveies set up the familial peculiarity of West and Cardinal African lions from other extant African populations. Interventions to salvage West African lions are desperately required. However explicating effectual preservation schemes has been hampered by a deficiency of informations on the species ' current distribution, position, and possible direction lacks of protected countries ( PAs ) harbouring lions. Our survey synthesized available adept sentiment and field informations to shut this cognition spread, and formulate recommendations for the preservation of West African lions. We undertook king of beasts studies in 13 big ( > 500 km2 ) PAs and compiled grounds of king of beasts presence/absence for a farther eight PAs. All PAs were situated within Lion Conservation Units, geographical units designated as precedence king of beasts countries by wildlife experts at a regional king of beasts preservation workshop in 2005. Lions were confirmed in merely 4 PAs, and our consequences suggest that merely 406 ( 273–605 ) lions remain in West Africa, stand foring < 250 mature persons. Confirmed king of beasts scope is estimated at 49,000 km2, or 1.1 % of historical scope in West Africa. PAs retaining lions were larger than PAs without lions and had significantly higher direction budgets. We encourage alteration of king of beasts taxonomy, to acknowledge the familial peculiarity of West African lions and highlight their potentially alone preservation value. Further, we call for listing of the king of beasts as critically endangered in West Africa, under standard C2a ( two ) for populations with < 250 mature persons. Finally, sing the comparative poorness of king of beasts scope provinces in West Africa, we call for pressing mobilisation of investing from the international community to help scope provinces to increase direction effectivity of PAs retaining lions.


The king of beasts ( Panthera Leo ) was the most successful big carnivore during the late Pleistocene, when the species ' scope extended from South Africa, across Eurasia, and into the southern United States. Today, the king of beasts 's scope is restricted to Africa and one population of the Asiatic sub-species, P. l. persica, in India. While the endangered Asiatic population is presently considered stable, lion populations in Africa are in diminution and the African sub-species, P. l. Leo, is considered vulnerable. Recent analyses established that the African king of beasts has lost at least 75 % of its original home ground, with fewer than 35,000 wild African lions staying. The chief drivers of king of beasts diminutions are large-scale habitat transition, prey base depletion through unsustainable hunting, and the retaliatory violent death of lions due to comprehend or existent human-lion struggle. The state of affairs is most critical in West Africa, where lions have been considered regionally endangered since 2004, and where < 500 persons may prevail.

West African lions represent a population with alone familial and preservation value. Recent molecular and morphological informations covering the species ' full historical scope suggests that lions in Central, West and North Africa ( the latter now extinct ) are distinguishable from lions in Eastern and Southern Africa and portion a common ascendant with lions in Asia – . These consequences set up that the chief subdivision of modern lions is within Africa, and question the current dichotomous division into an African and an Asiatic sub-species ; a division which is still widely supported, including by the IUCN Red List. Furthermore, they demonstrate that lions in West Africa contain mtDNA haplotypes non found in other king of beasts populations, promoting the preservation significance of the few staying West African populations, .

Conservation intercessions to salvage these populations are now desperately required. However, explicating intercessions is limited by few field informations on the species ' current distribution, copiousness, and prevailing drivers of diminutions in West Africa. While lions have been the object of extended research attempt in parts of Eastern and Southern Africa, they have been mostly ignored in West Africa. Of 463 articles on African lions in the ISI Web of Science™ ( Thompson Scientific ) in 2005, non one focused specifically on lions in West Africa. To turn to this shortage, we foremost collated available informations and adept sentiments on king of beasts distribution and position in West Africa. We so undertook field studies in 13 big ( > 500 km2 ) protected countries ( PAs ) where lions were reported during this procedure, to find king of beasts presence/absence and estimation king of beasts population size. For a farther eight big PAs suspected to harbour lions, we compiled field study informations from the literature and via interviews. Using standardised rating toolkits for protected country direction effectivity, we compared current direction public presentation of PAs known or suspected to harbour lions and those from which lions have likely been extirpated. With these informations, we provide a comprehensive rating of the tenuous position of the West African king of beasts, and do recommendations for the taxa 's preservation.

Study country

We restrict our analysis to West Africa, as defined by the United Nations geoscheme ( hypertext transfer protocol: //unstats.un.org/unsd/methods/m49/m49regin.htm ) , including all states from Senegal to Nigeria ( Fig. 1 ) . The same categorization was used for the listing of the king of beasts as regionally endangered in West Africa. Historically, lions occurred in all biomes in West Africa, with the exclusion of the coastal Upper and Lower Guinean Forests and the inside of the Saharan Desert ( Fig. 1 ) . The prostration of king of beasts scope in West Africa is ill documented, but appears to be linked to large-scale habitat loss outside PAs through transition to agribusiness. Consequently, king of beasts scope in this part is mostly restricted to PAs, . While several West African states have big PAs, mean PA public presentation in West Africa ranked poorest in a cross-continental comparing: big mammal populations in eleven West African PAs declined by an norm of 85 % between 1970–2005, compared to an mean 59 % diminution across the continent. Population prostration within these PAs appears to be driven by commercial bushmeat development, providing local markets in West Africa – , and has contributed to conveying several iconic big mammal species to the threshold of extinction in their West African scope, .

Consultation of adept sentiment on current king of beasts scope

In 2005, the IUCN and the Wildlife Conservation Society ( WCS ) organized a king of beasts preservation workshop for wildlife governments from all king of beasts scope states within West and Central Africa. The workshop consisted of a proficient session to map current king of beasts scope and position, followed by a strategic planning session to develop lion preservation schemes. The proficient session was modeled after the Range Wide Priority Setting procedure developed by WCS for panthers. Experts were guided to bring forth maps of current king of beasts scope and delimit countries harbouring known or suspected populations called Lion Conservation Units ( LCUs ) . LCU boundary line relied on believable grounds of king of beasts presence within the predating 10 old ages, and for each LCU participants assigned lion population tendencies and approximative king of beasts population size. Although the informations presented at the 2005 workshop now day of the month from 1995 onwards, at the oncoming of our field studies in 2006 ( see below ) , it represented the most comprehensive and dependable dataset on king of beasts distribution in West Africa.

Field studies

Between October 2006 and May 2012, we conducted field studies in PAs within designated LCUs, to 1 ) confirm king of beasts presence for PAs where grounds of king of beasts happening was missing, and 2 ) set up king of beasts population estimations for PAs where lions occurred. Although designated LCUs included both protected and non-protected countries, lions are mostly absent outside PAs in West Africa, . Consequently, we restricted fieldwork to PAs ( IUCN PA classs I–VI ) within LCUs ( henceforth: LCU PAs ) , including two PAs expecting formal appellation ( Table 1 ) . A primary determiner of king of beasts extinction hazard within a given PA is its size, and we hence restricted our study attempt to big ( > 500 km2 ) LCU PAs, based on findings on critical PA size for lions from anterior surveies, . In LCUs with multiple PAs, we concentrated study attempts in the largest Pas with the highest protection position ( harmonizing to IUCN PA classs ) .

Establishing king of beasts presence/absence

Survey methods normally used for African Savannah mammals, such as aerial studies or line transects, typically yield few observations of big tellurian carnivores. Consequently, anterior attempts to set up big carnivore happening and/or copiousness over big spacial graduated tables relied on interviews, remote cameras, or path studies. We preponderantly employed path studies, owing to their relatively high sensing efficiency, and low attempt and cost. All studies squads included experient perceivers, and we ascribed field mark to species based on pugmark features. Occasionally teams found ambiguous paths, chiefly regionally rare species such as African wild Canis familiaris ( Lycaon pictus ) and cheetah ( Acinonyx jubatus ) . In such instances we documented pugmarks photographically, and presented exposures to a panel of experts for species confirmation. We omitted a little figure of records from analysis if they could non be assigned to species unambiguously. In PAs with an integral web of soil paths, we conducted vehicle-based path studies, with two perceivers seated on the bull-bar of a vehicle driven at a maximal velocity of 10–20 kilometers h−1. In PAs without penetrable roads, we conducted track hunts on pes along approximately predefined study circuits, following game trails, dry river bottoms, abandoned soil paths or other additive characteristics normally used as travel paths by lions and other big carnivores. These circuits incorporated home ground characteristics that could be expected to pull larger herbivores, such as H2O reservoirs, flood plains, saltlicks and fens, or other sites with high herbivore copiousness indicated by PA staff. In Mole NP and Gbele Resource Reserve in Ghana, camera traps were the primary study method. At those PAs, we deployed DeerCam DC-300 ( Non Typical, Wisconsin, USA ) camera units at ∼1 kilometer intervals, aiming, as with pes studies, characteristics expected to maximise king of beasts gaining control chance. In Mole NP, we concentrated pin downing attempt in the cardinal and southeasterly parts known to incorporate higher quarry densenesss and cardinal dry-season H2O beginnings. In Comoé National Park ( NP ) in Côte d'Ivoire, West Africa 's largest Neptunium at > 10,000 km2, we conducted an aerial study prior to our land study, to place countries with of import concentrations of possible king of beasts quarry. We restricted the resulting pes study to those countries.

Estimating king of beasts population size

If lions were found in a given PA, we used systematic path counts or call-ups to gauge king of beasts population size, depending on local conditions. Neither method has of all time been locally calibrated in West Africa. However, we preferred path counts due to the by and large consistent relationship between king of beasts path densenesss and existent king of beasts densenesss observed across a broad scope of different climatic and habitat-related conditions in Eastern and Southern Africa. One of import caution of this method is that the relationship between path densenesss and existent densenesss varies with substrate type, and we hence recorded substrate type every 500 metres along spoor transects, to enable us to choose the appropriate relationship from Funston et Al. In two PAs with king of beasts presence where the hapless province of the route web precluded the usage of vehicle-based path counts, we conducted call-ups to gauge king of beasts population size. The call-up method requires standardization experiments to measure local response distance and response rate of lions to broadcasts. Because king of beasts observations were highly rare during our studies, we obtained merely one response distance estimation in heavy Sudano-Sahelian forest ; one male king of beasts encountered opportunistically 2.5 kilometer from a call-up station, was observed once more at the station 26 proceedingss after the oncoming of the broadcast, 50 proceedingss after the initial observation. As the male was going in the opposite way when foremost seen, we consider this a conservative estimation for a maximal response distance in dense forest, and used this value here. To derive probationary estimations of king of beasts population denseness based on our call-up consequences, we used the scope of published figures on king of beasts response rates, . Study design and information analysis of our systematic path counts followed Funston et al. , while our protocol for call-ups followed Ferreira and Funston. All field studies were carried out in close coaction with the several national wildlife governments, and involved senior PA research staff. Our study work was hence considered an built-in portion of preexisting PA monitoring activities, and wildlife governments waived demands for formal research clearance and PA entry fees. All field methods used were wholly noninvasive and did non necessitate the handling or sampling of unrecorded animate beings, and our study work did hence non necessitate blessing from an ethics commission.

Measuring direction of LCU PAs

Prior surveies highlighted the strong impact of PA features ( e.g. PA size ) , human population denseness at PA borders, and PA direction variables ( e.g. PA direction budgets ) on king of beasts continuity and population position inside PAs, , . Therefore, we investigated correlativities between king of beasts continuity and a figure of uninterrupted PA variables, including one-year budget, staff figure, country, environing human population denseness and IUCN direction class, utilizing univariate analyses ( Analysis of Variance ( ANOVA ) and Mann Whitney U trials ) where sample size allowed. We tested for univariate correlativities between the PA variables utilizing univariate additive arrested development or spearman 's rank. We estimated human population denseness within a 5 kilometer buffer around each PA based on human population informations from the AfriPop Project ( www.afripop.org ) , utilizing PA lineations obtained from the World Database of Protected Areas ( hypertext transfer protocol: //protectedplanet.net/ ) . We extracted uninterrupted direction variables from WWF/World Bank Management Effectiveness Tracking Tool ( METT ) appraisals carried out in the part. The METT is one of the most widely used appraisal tools for Protected Area Management Effectiveness ( PAME ) , and is designed to be completed by PA directors, staff and stakeholders. Besides provisioning of uninterrupted PA direction variables such as budgets and staff Numberss, the methodological analysis encompasses a rapid appraisal based on a scorecard questionnaire of 30 inquiries, with an ordinal four-point graduated table ( 0–3, with 3 stand foring the best direction scenario ) . The complete METT questionnaire templet is provided in Supporting Information S1, and Table S1 summarizes the 13 METT scored inquiries included in our analysis ; we selected direction facets probably to act upon the ability of a PA to implement ordinances and cut down hunting force per unit area, to supply penetration into the directors ' perceptual experience of current PA public presentation ) . A comprehensive list of all PA direction variables used in our analyses, variable birthplace, and matching sample sizes can be found in Table S2. All statistical analyses were performed in R.

Lion presence and absence in West African PAs

Wildlife experts go toing the 2005 workshop identified 17 LCUs in West Africa ( Fig. 1 ) , numbering 254,430 km2, or 5.8 % of historic king of beasts scope in West Africa. We identified 21 big ( > 500 km2 ) PAs within those LCUs ( Fig. 1, Table 1 ) , with a entire country of ∼95,000 km2, or 37 % of entire LCU extent. We surveyed 13 of those PAs for lions, while the staying PAs were surveyed by other research workers concentrating on lions ( n = 2 ) , Western Pan troglodytess Pan troglodytes ssp. verus ( n = 2 ) , and general faunal stock lists ( n = 4 ) ( Table 1 ) . Of the 21 LCU PAs surveyed, lions were confirmed in merely four ( Table 1 ; Fig. 2 ) . In two extra PAs, both in Guinea, lions had non been observed for > 10 old ages ; nevertheless, believable studies of voices suggest they may still be present. Among the four PAs in which king of beasts continuity was confirmed, three contain < 50 persons, and the lone big population is in the W-Arly-Pendjari ( WAP ) , with an estimated 356 ( scope: 246–466 ) lions ( Table 1 ) . The entire figure of lions staying in West Africa is estimated at 406 ( scope: 250–587 ) persons, while the confirmed king of beasts scope ( the sum size of PAs where lions were confirmed, including possible sites in Guinea ) is estimated at 49,000 km2, or 1.1 % of historic king of beasts scope in West Africa.

Management of LCU Protected Areas

( A ) PAs where lions are confirmed present or are potentially still present ( A ) ; ( B ) PAs where lions were considered absent. The missive W represents tonss for W-Arly-Pendjari while M indicates those for Mole NP ( see text ) . Management tonss range from 0–3, with 3 stand foring the best direction scenario. For illustration, in the instance of ‘Current Budget’ 0 = No PA budget ; 1 = inadequate budget which creates serious direction restraints ; 2 = acceptable budget, but could be farther improved to to the full accomplish effectual direction ; 3 = sufficient budget which to the full meets the demands of the PA. See Table S1 for full descriptions of tonss.


The king of beasts has undergone a ruinous prostration in West Africa. Our consequences suggest that lions have lost about 99 % of their historic scope, and that merely ca. 400 single lions persist across the part. Most of these lions ( ca. 350 persons, or 88 % of the entire population ) persist in a individual population in WAP, and there is strong grounds for ongoing diminutions in the part 's other three populations. In Nigeria, Numberss dropped from an estimated 44 lions in 2009 to 34 in 2011. In Senegal 's Niokolo-Koba NP, go oning black diminutions in prey populations ( Fig. 5B ) are about surely doing attendant diminutions in lions. These tendencies suggest that WAP already or will shortly incorporate > 90 % of West African lions. Given that 40–60 % of a king of beasts population typically consists of immature persons, , and that our path counts in WAP included big greenhorns and sub-adults, it is really likely there are < 250 grownups staying in the full West African part. Consequently, our consequences warrant listing of the king of beasts as critically endangered in West Africa under standard C2a ( two ) , which applies for worsening populations with < 250 mature persons, where > 90 % of persons persist in one subpopulation.

Precedences for king of beasts preservation in West Africa

Our studies covered all big ( > 500 km2 ) , officially designated PAs within LCUs in West Africa. While king of beasts scope in this part is mostly restricted to PAs, , we can non except the possibility that some lions roam outside the surveyed PAs. However, the 21 LCU PAs covered in this survey represent the best staying king of beasts home ground in West Africa. We deem study attempt adequate ( see Table S3 ) to pull illation on the happening of resident lions across sites, and we are confident that no resident king of beasts populations were overlooked by our attempts. Further study work may be required in Haut Niger NP and Kankan FR in Guinea to measure the possible presence of lions. However, given the deficiency of physical grounds for over a decennary and the hapless direction tonss of those two PAs ( see Table S4 and below ) , we believe any staying populations would be relict and near to extinction. While continued study and monitoring work is warranted, the highest preservation precedence for lions in the part should be beef uping protection of the known staying populations.

Lions are more likely to vanish from little PAs than from larger 1s. Critical PA size for lions based on informations from East Africa is 291 km2, utilizing an mean king of beasts denseness of 16.2 grownups 100 km−2. In comparing, mean denseness across our four sites with lions was 1.0 lions 100 km−2, > 15 times lower than in East Africa. Assuming the same lower bound for a feasible king of beasts population size in West Africa, critical PA size would transcend 4,000 km2 at current king of beasts densenesss and even that may be unequal. Newmark revealed that extinction rates in Ghanese PAs were estimated to be 13–77 times higher than in equivalent-sized PAs in Tanzania, proposing that larger size entirely a may supply deficient protection against the intense hunting force per unit area impacting West African PAs. In add-on to the lower transporting capacity of West African savannas for big herbivores, higher extinction hazards for West African mammals is driven by intense bushmeat runing force per unit area within and next to PAs, , facilitated by uneffective PA direction ( Fig. 4 ) . Our findings highlight the pressing demand for really big ( > 4,000 km2 ) and good protected PAs to guarantee the endurance of lions and other threatened big mammals in West Africa. Three of four extant king of beasts populations in West Africa occur in PAs stopping point to or larger than 4,000 km2 ( Table 1 ) , stand foring the best chances for salvaging the taxon.

WAP presently harbors the lone population > 50 animate beings, and is the most feasible. However, lion population denseness is highly low in the eastern half of WAP, i.e. the tri-national W NP ( Henschel et al. in homework ) . An aerial study covering W instantly following our king of beasts study in 2012, recorded > 50,000 caput of cowss inside the national park, underscoring the weak direction effectivity in W NP. In contrast, the western half, Arly-Pendjari, supports higher king of beasts densenesss, stable or increasing prey populations ( Fig. 5A ) , and incursions by farm animal into the PA are rare.The blunt contrast in direction effectivity between the eastern and western halves of WAP may be due to the disparity in direction budget allotments ; of the US $ 197/km2 available for the WAP in sum, 90/km2 are spent in W, compared to 323/km2 in Arly-Pendjari ( Henschel et al. in homework ) . A significantly higher operational budget is required in W to achieve preservation outcomes comparable to Arly-Pendjari.

Lion populations in Niokolo-Koba and Kainji Lake NPs are little and appear to be worsening. While no informations on direction effectivity and king of beasts quarry populations exist for Kainji Lake NP, direction effectivity tonss are low in Niokolo-Koba NP, potentially due to inadequate support ( Table S4 ) , and prey populations have collapsed to highly low degrees over the past 20 old ages ( Fig. 5B ) . Both PAs hold great possible due to their big size, and are surrounded by suited king of beasts home ground and moderate human population densenesss ( Table S3 ) . Furthermore, Kainji Lake NP is potentially still connected to WAP, through suited king of beasts home ground in Benin. Besides lions, Niokolo-Koba besides harbors the last of import population of the critically endangered Western elephantine eland ( Tragelaphus derbianus ssp. derbianus ) , and the merely confirmed population of the critically endangered West African sub-population of African wild Canis familiariss ( Table S3 ) . Both PAs will necessitate immediate fiscal and proficient aid to debar the local ablation of lions and other critically endangered taxa.

At 2,244 km2 Nigeria 's Yankari Game Reserve is smaller than our putative lower limit and its king of beasts population is really little and worsening ( Table 1 ) . Yankari is wholly surrounded by intensive cultivation, and the second-highest human population denseness of all 21 LCU PAs surveyed ( Table S3 ) . As a effect, Yankari 's lions and so all large-medium mammals are likely to be efficaciously isolated from neighbouring populations in Nigeria ( Kainji Lake NP, ca 650 kilometers distant ) and Cameroon ( Benoué Complex, ca 260 kilometers distant ) . Drastic intercessions, such as fencing the modesty, may be the lone solution to safeguard this population. Fencing Yankari could forestall inevitable invasion by people and farm animal, cut down human-lion struggles at the PA boundary and possibly cut down incursion of the PA by poachers.

The province of PAs in West Africa

Of 12 PAs with direction appraisals, six had no budget for direction activities, and where budgets existed they were much lower than required to conserve king of beasts populations efficaciously. PAs with confirmed or likely king of beasts presence had larger budgets than those with lions absent, and WAP had both the largest king of beasts population and the highest one-year budget, at US $ 197/km2 ( Table S4 ) . WAP moreover systematically received among the highest tonss for direction effectivity of all PAs harbouring lions ( Fig. 4B ) . For PAs where information on population tendencies of chief king of beasts quarry species were available, WAP ( represented through Pendjari NP, where one-year wildlife counts are conducted ) was besides the lone site where wildlife Numberss were stable or increasing ( Fig. 5 ) . As a farther indicant of preservation success in WAP, this site seaports by far the largest staying elephant population in West Africa, and one of the last leftover populations of the critically endangered Northwest African chetah ( Acinonyx jubatus ssp. hecki ) . Even so, WAP 's budget is an order of magnitude below the estimated > US $ 2000/km2 budget required to keep lions in unfenced PAs, proposing that WAPs success may non be sustainable and will necessitate increased support in future.

Staff Numberss for LCU PAs were by and large low, changing between 0–4 staff per 100 km2. Many PAs reported that staff wages were paid straight by national authorities and non through the single PA direction budget. Where staff are paid by cardinal authorities, yet few or no financess are available for active direction of PAs, the reported staff Numberss likely overestimate preservation attempt: nine of the 12 PAs assessed utilizing METT reported holding either no jurisprudence enforcement activity at all ( four PAs ) , or major lacks in staff capacity/resources to consequence patrols ( five PAs ) . Tax assessors for Comoé NP, which has 54 patrol staff, commented that “Staffing is really low for the size of the park ; merely 2 of 5 sectors of the park are operational, with 1 vehicle, limited staff, no equipment and deficiency of training” . In Haut Niger, which has 15 patrol staff, assessors noted that in pattern, agents do no or really few patrols due to a deficiency of resources and motive ; as a consequence poaching and illegal logging is widespread. Brugière notes that bing PAs in Guinea are basically paper-parks, i.e. they have no staff, direction program or operating budget. Consequently, even in the largest officially gazetted PA in Guinea, the Kankan Faunal Reserve, where lions are potentially still present, ca 20,000 people live within the PA, poaching force per unit area is high, and antelope population densenesss are highly low.

WAP represents the last fastness for lions in West Africa. Conservation intercessions in WAP are to a great extent subsidized by big international support organic structures, such as the European Union and the World Bank. However, overall investing in preservation activities is highly low in West Africa, compared to Central, Eastern and Southern Africa. Sing that all eleven former or current king of beasts scope states in West Africa are among the 50 poorest states in the universe, and that six are classified as least developed states ( informations from World Bank ; hypertext transfer protocol: //databank.worldbank.org/data/home.aspx ) , king of beasts scope provinces in West Africa will be unable to mobilise the resources required to procure their staying king of beasts populations. That will trust on the proviso of significant fiscal and proficient aid to run provinces, chiefly by the international community, to increase direction effectivity of PAs with lions. For any such investings, it will be imperative that a ) preservation initiatives assure sound administration over the financess, and B ) adequate support degrees are sustained in the long-run to accomplish desired outcomes ; a reappraisal of best patterns can be found in Blom et al..

It is imperative to turn to really widespread poaching of king of beasts quarry species and illegal violent death of lions by pastoralists within and around PAs, , . We believe pressing precedence must be given to a double scheme that focuses on 1. increasing the Numberss, expertness, and runing budgets of enforcement forces in PAs with lions, to control the violent death of king of beasts quarry and illegal incursions into PAs by pastoralists, and 2. cut downing human-lion struggle in affected communities surrounding PAs, by uniting improved husbandry patterns with community sensitisation, to cut down farm animal losingss to marauders and better local negative perceptual experiences of big carnivores. Investing should besides be directed toward developing and heightening photographic touristry in politically stable states such as Benin and Senegal. This will assist to make and keep economic inducements for king of beasts preservation, and develop digesting gross watercourses for PA direction non entirely reliant on giver support.

Conclusions & Recommendations

Sing the comparative poorness of king of beasts scope provinces in West Africa, we call for the mobilisation of significant and pressing investing by the international community to help these states in bettering direction effectivity of PAs incorporating lions. Lions persist in some of the largest and most integral protected landscapes in West Africa, where they co-occur with some of the last leftover populations of critically endangered mammals including Northwest African chetah, Western elephantine elands and African wild Canis familiariss. Further impairment of those last wilderness countries in West Africa will probably do the loss of genetically distinguishable populations of magnetic megafauna and farther preclude already tenuous, possible hereafter gross watercourses from photographic touristry for West African states. Without immediate action, we believe the chance to salvage both will be lost.


We thank the several wildlife governments for helping attempts to carry on king of beasts studies in the protected countries covered in the present survey. Specific thanks go to M. Sidibé and I. Ndao in Senegal, A. Gonto, D. Azani and I. Herbinger from the Wild Chimpanzee Foundation in Côte d'Ivoire, M. Sam in Ghana, A. Tehou, M. Kiki and C. Sewade in Benin, P. Bouché in Burkina Faso, H. GARBA in Niger, every bit good as G. Nyanganji from WCS in Nigeria. We thank L. Petracca ( Panthera ) for aid with GIS analyses, F. Leverington for the designation of extra PA direction informations, and C. Breitenmoser, one anon. referee and the editor for helpful remarks on the manuscript.

Head Coach more

Caldwell joined Indianapolis from Tampa Bay, where he served in the same capacity during the 2001 season. Caldwell has more than 20 old ages of collegial experience. Caldwell spent 1993-2000 as caput manager at Wake Forest. In 1999, Caldwell led the school to its first winning season and bowl game since 1992. Wake Forest defeated Arizona State, 23-3, in the Aloha Bowl. During Caldwell 's term of office, Wake Forest ranked among the state 's Top 25 squads in go throughing discourtesy on four different occasions, including 10th in the NCAA in 1995. That season, signal caller Rusty LaRue established seven NCAA passing records.

Caldwell served as an adjunct manager at Southern Illinois ( 1978-80 ) , Northwestern ( 1981 ) , Colorado ( 1982-84 ) , Louisville ( 1985 ) and Penn State ( 1986-92 ) . Caldwell joined Penn State as broad receiving systems coach. He so coached signal callers the undermentioned season and added passing game coordinator duties in 1988. At Penn State, Caldwell tutored signal caller Kerry Collins, who went on to win the Davey O'Brien Award as the state 's top college signal caller and the Maxwell Award as the state 's most outstanding participant. Caldwell has coached in six bowl games and won a national title with Penn State in 1986. In add-on to functioning on Joe Paterno 's staff that won a national title, Caldwell tutored under three other managers who won collegiate rubrics ( Rey Dempsey, Southern Illinois ; Bill McCartney, Colorado ; Howard Schnellenberger, Louisville ) .


Lions are besides the lone cats that live in big, societal groups called “prides.” A pride can hold 3 to 30 lions and is made up of lionesses ( female parents, sisters, and cousins ) , and their greenhorn, along with a few unrelated grownup males. The pride has a close bond and is non likely to accept a alien. The unrelated males stay a few months or a few old ages, but the older lionesses stay together for life. In dry countries with less nutrient, prides are smaller, with two lionesses in charge. In home grounds with more nutrient and H2O, prides can hold four to six grownup lionesses. Both males and females scent grade to specify their district.


The San Diego Zoo Safari Park has had lions on exhibit since it opened in 1972. In 2004, the Safari Park’s Lion Camp opened with six endearing six-month-old Transvaal king of beasts greenhorns freshly arrived from a installation in Africa. Lion Camp looks like a spot of African home ground, so guests get to see tonss of natural king of beasts behaviours, watching the cats as they romp in the grass, explore the logs and stones, or sit and watch the antelope, camelopard, and rhinos in the nearby African Plains field enclosure. Guests can come right up to the big glass panels for some eye-to-eye minutes with the animals. There is even a campaign vehicle, a favourite resting topographic point for our pride. You may see a king of beasts perched on its roof and detecting life from the driver’s place!

Two of those original six Lion Camp occupants, M’bari and Etosha, now hold tribunal in the Zoo’s Elephant Odyssey, an country of the Zoo where we spotlight the animate beings that lived here in Southern California over 10,000 old ages ago with their contemporary opposite numbers. Here you’ll find a lifesize statue of the American king of beasts, which lived here 12,000 old ages ago. Our lions study district that includes a foothill environment with bouldery inclines, trees, grasses, and a watercourse. Specially het stones make the perfect lounging topographic point for the male monarch of animals. At this “kingdom, ” invitees get near plenty to number the cats’ hair's-breadth musca volitanss and bask in the lions’ aureate regard. M’bari awes visitants with his olympian bearing and late-afternoon turns of boom.

Life for our lions is filled with new and unexpected experiences, merely like it would be for them in the natural state ; nevertheless, it 's up to their keepers to supply the large cats with those experiences by offering them a assortment of things to whiff, gustatory sensation, or drama with. We call this enrichment. Keepers frequently place interesting aromas in their exhibit, found in such points as wood shaves from our barns, herbs like cloves or cinnamon rubbed on a stone, or fox urine sprayed on a wall. Food enrichment can include big, blocks of ice, like a immense ice lolly, filled with balls of meat, called “meat-sicles, ” or whole fish—a “fish-sicle” ! Cardboard boxes, palm fronds, and provender pokes make great playthings, excessively. And the lions love to play with big, heavy-duty plastic balls, turn overing, fliping, and even swooping on them, wholly in good merriment. Thankss to their keepers and trainers, there is ne'er a dull twenty-four hours.


Are lions in problem? Due to many issues such as disease, runing by worlds, and loss of home ground, the population of lions in the natural state is going really refering to environmentalists. Natural home ground for lions is now found merely in protected militias, and king of beasts motion between prides is going more limited. While king of beasts hunting is banned in many African states, trophy hunting is still allowed in some topographic points. In other countries, there are so many lions for so small infinite that Texas Rangers frequently put the females on birth control to cut down the figure of cubs born. Ranchers sometimes poison lions that prey on farm animal.

San Diego Zoo Global supports king of beasts preservation. We provide financess to two organisations in Africa that work to assist lions and other wildlife: the Northern Rangelands Trust and the Grevy’s Zebra Trust. The preservation of lions has to be reconciled with the demands of worlds. Some struggle may be ineluctable in countries where agribusiness or farm animal farming compete with cat home grounds, but it can be minimized, and local people must experience that attempts are being made to protect their involvements. Education is a cardinal constituent of preservation, supplying information to all degrees of the community about the function of cats in their home ground and ways to conserve them.

Michael Rothstein ESPN Staff Writer

The Lions picked up TE Eric Ebron 's fifth-year option this forenoon. On 97.1 FM in Detroit, Quinn explained why the procedure took so long: `` I think that 's portion of my occupation, to maintain every possible option unfastened on any portion of this organisation. The bill of exchange is evidently a large event to add endowment to the squad so I think the conference regulations put that May 2 day of the month in there for a ground, it 's non merely an arbitrary twenty-four hours. I think they do that a twosome yearss after the bill of exchange for a ground, to give squads clip to measure what they get in the bill of exchange and so do a determination on their former first unit of ammunition choices. This is something we 've thought about, talked about internally here evidently since the terminal of the season and I sort of had my head made up likely last hebdomad at some point but I wanted to acquire through the bill of exchange and see. You ne'er know what can go on through the bill of exchange and what sort of participants could steal to you. ''


The seasoned defensive tackle wanted to do certain he was wholly healthy heading into the new season and even underwent neurological test to find if he has sustained any encephalon harm over his calling. The consequences showed no encephalon harm, harmonizing to the defensive tackle. `` The encephalon is good to go.I wanted to do certain my caput was good. He ( the brain doctor ) said everything was reasonably good - truly good, actually.Just to do certain, with all things that are traveling on with the encephalon and material throughout the conference, I do n't desire to hold jobs when I 'm older '' , Ngata said. The defensive tackle missed some clip last twelvemonth due to calf and quad issues but still managed to enter 22 tackles and 1.5 pokes through 13 games.

The organisation was on the fencing about perpetrating to Ebron through 2018 after a largely dissatisfactory first two seasons in the conference, but the Lions seemingly saw adequate growing in his public presentation in 2016 to warrant maintaining him in the crease a small thirster. Aside from adding Michael Roberts in the 4th unit of ammunition of last weekend 's NFL bill of exchange, the Lions have n't made many major add-ons at tight terminal during the offseason, so Ebron one time once more looks primed to open the approaching run as the squad 's starting motor. His 2016 sums of 61 gimmicks and 711 paces were nice, but he 'll necessitate to promote his touchdown sum ( one ) if he hopes to go an upper-tier phantasy tight terminal.

Lion News

Females normally stay in their female parents ' prides for life, unless nutrient scarceness forces them out. Young males are driven from their prides when they grow big plenty to vie with the dominant males ( normally between the ages of two and four ) . Young males join in alliances, normally with brothers and cousins, and hunt for a pride to take over. Males come ining a new pride will kill all greenhorns that can non run from them. Adult males that are fortunate plenty achieve residence within a pride clasp term of office for an norm of two old ages, frequently go forthing due to eviction by another alliance of males. In India, female and male lions live apart, fall ining merely to copulate.

More than 10,000 old ages ago, lions thrived from North and South America to Europe, Africa and Asia. Today, following climatic alterations and after centuries of runing and habitat debasement by people, lions live in scattered home grounds across Africa ( with the exclusion of the Gir Forest lions, which live in a national park in northwest India ) . Within these countries, lions still face dangers, including habitat loss and runing. Many have died from diseases such as distemper, which is spread by domestic Canis familiariss from small towns near natural home ground. To maintain lions from going every bit rare as Liberation Tigers of Tamil Eelams, big sweeps of home ground must be carefully protected. In disease-ridden countries, runing Canis familiariss must be kept off from lions or immunized.


Lions are the lone genuinely societal cats, with related females populating together in prides overseen by male alliances that compete for ownership in fierce and frequently fatal conflicts. These brilliant animals are found in sub-Saharan Africa, and there is besides a little stray population of Asian lions to be found in the Gir Forest of western India. Lions are marauding carnivores and it 's the females that perform most of the hunting at dark ; the bulk of the quarry being antelope, zebra and wildebeest. They breed year-round and it’s estimated that lions copulate around 3,000 times for every greenhorn that is born and survives for over a twelvemonth. Did you cognize? Lions are the lone cats with a mane.


The king of beasts ( Panthera Leo ) is one of the large cats in the genus Panthera and a member of the household Felidae. The normally used term African king of beasts jointly denotes the several races in Africa. With some males transcending 250 kilogram ( 550 pound ) in weight, it is the second-largest life cat after the tiger, excluding loanblends like the liger. Wild lions presently exist in sub-Saharan Africa and in India ( where an endangered remnant population resides in and around Gir Forest National Park ) . In ancient historic times, their scope was in most of Africa, including North Africa, and across Eurasia from Greece and southeasterly Europe to India. In the late Pleistocene, about 10,000 old ages ago, the king of beasts was the most widespread big land mammal after worlds: Panthera leo spelaea lived in northern and western Europe and Panthera leo atrox lived in the Americas from the Yukon to Peru. The king of beasts is classified as a vulnerable species by the International Union for Conservation of Nature ( IUCN ) , holding seen a major population diminution in its African scope of 30–50 % over two decennaries during the 2nd half of the 20th century. Lion populations are indefensible outside designated militias and national Parkss. Although the cause of the diminution is non to the full understood, habitat loss and struggles with worlds are the greatest causes of concern. Within Africa, the West African king of beasts population is peculiarly endangered.

In the wilderness, males rarely live longer than 10 to 14 old ages, as hurts sustained from continual contending with rival males greatly cut down their length of service. In imprisonment they can populate more than 20 old ages. They typically inhabit savanna and grassland, although they may take to bush and forest. Lions are remarkably societal compared to other cats. A pride of lions consists of related females and offspring and a little figure of grownup males. Groups of female lions typically hunt together, feeding largely on big hoofed mammals. Lions are apex and keystone marauders, although they are besides adept scavengers obtaining over 50 per centum of their nutrient by scavenging as chance allows. While lions do non typically hunt worlds, some have. Sleeping chiefly during the twenty-four hours, lions are active chiefly at dark ( nocturnal ) , although sometimes at twilight ( crepuscular ) .

Highly typical, the male king of beasts is easy recognised by its mane, and its face is one of the most widely recognised animate being symbols in human civilization. Word pictures have existed from the Upper Paleolithic period, with carvings and pictures from the Lascaux and Chauvet Caves in France dated to 17,000 old ages ago, through virtually all antediluvian and medieval civilizations where they one time occurred. It has been extensively depicted in sculptures, in pictures, on national flags, and in modern-day movies and literature. Lions have been kept in menageries since the clip of the Roman Empire, and have been a cardinal species sought for exhibition in menagerie over the universe since the late 18th century. Zoos are collaborating worldwide in engendering plans for the endangered Asiatic races.

Taxonomy and development

The king of beasts 's closest relations are the other species of the genus Panthera: the tiger, the snow leopard, the panther, and the leopard. Surveies from 2006 and 2009 concluded that the panther is a sister species to the king of beasts and the leopard is a sister taxon to the jaguar/lion clade while 2010 and 2011 surveies have swapped the places leopard and panther. P. leo evolved in Africa between 1 million and 800,000 old ages ago, before distributing throughout the Holarctic part. It appeared in the dodo record in Europe for the first clip 700,000 old ages ago with the races Panthera Leo fossilis at Isernia in Italy. From this king of beasts derived the ulterior cave king of beasts ( Panthera leo spelaea ) , which appeared about 300,000 old ages ago. Lions died out in northern Eurasia at the terminal of the last glaciation, about 10,000 old ages ago ; this may hold been secondary to the extinction of Pleistocene megafauna.


Traditionally, 12 recent races of king of beasts were recognised, distinguished by mane visual aspect, size, and distribution. Because these features are really undistinguished and demo a high single variableness, most of these signifiers were likely non true races, particularly as they were frequently based upon zoo stuff of unknown beginning that may hold had `` dramatic, but unnatural '' morphological features. Today, merely eight races are normally accepted, although one of these, the Cape king of beasts, once described as Panthera Leo melanochaita, is likely invalid. Even the staying seven races might be excessively many. While the position of the Asiatic king of beasts ( P. l. persica ) as a race is by and large accepted, the systematic relationships among African lions are still non wholly resolved. Mitochondrial fluctuation in living African lions seemed to be modest harmonizing to some newer surveies ; hence, all sub-Saharan lions have sometimes been considered a individual races. However, a recent survey revealed lions from western and cardinal Africa differ genetically from lions of southern or eastern Africa. Harmonizing to this survey, West-central African lions are more closely related to North African and Asiatic lions than to Southern or Eastern African lions. These findings might be explained by a late Pleistocene extinction event of lions in western and cardinal Africa, and a subsequent recolonisation of these parts from Asia.

Previous surveies, which were focused chiefly on lions from eastern and southern parts of Africa, already showed these can be perchance divided in two chief clades: one to the West of the Great Rift Valley and the other to the E. Lions from Tsavo in eastern Kenya are much closer genetically to lions in Transvaal ( South Africa ) , than to those in the Aberdare Range in western Kenya. Another survey revealed there are three major types of lions, one North African–Asian, one southern African and one in-between African. Conversely, Per Christiansen found that utilizing skull morphology allowed him to place the races krugeri, nubica, persica, and senegalensis, while there was convergence between bleyenberghi with senegalensis and krugeri. The Asian king of beasts persica was the most typical, and the Cape king of beasts had features allying it more with P. l. persica than the other sub-Saharan lions. He had analysed 58 king of beasts skulls in three European museums.

Based on recent familial surveies, the Cat Classification Task Force of the IUCN SSC Cat Specialist Group has provisionally proposed to delegate the lions happening in Asia ( P. l. persica ) and West, Central and North Africa ( P. l. senegalensis, and P. l. Leo ) to the races Panthera Leo Leo, and the lions populating Southern ( P. l. bleyenberghi and P. l. krugeri ) and Eastern Africa ( P. l. azandica and P. l. nubica ) to the races Panthera Leo melanochaita. The U.S. Fish and Wildlife Service has followed this revised systematic categorization, as being based on `` the best available scientific and commercial information '' , in naming these two races as, severally, endangered and threatened.


Of the life, non-hybrid felines, the king of beasts is 2nd merely to the tiger in length and weight. Its skull is really similar to that of the tiger, although the frontal part is normally more down and flattened, with a somewhat shorter postorbital part and broader nasal gaps than that of a tiger. However, due to the sum of skull fluctuation in the two species, normally merely the construction of the lower jaw can be used as a dependable index of species. Lion color varies from light fan to yellowish, reddish, or dark ochraceous brown. The underparts are by and large lighter and the tail tussock is black. Lion greenhorns are born with brown rosettes ( musca volitanss ) on their organic structure, instead like those of a leopard. Although these fade as lions reach maturity, weak musca volitanss frequently may still be seen on the legs and underparts, peculiarly on lionesses.

Lions are the lone members of the cat household to expose obvious sexual dimorphism – that is, males and females look clearly different. They besides have specialised functions that each sex plays in the pride. For case, the lioness ( the huntsman ) lacks the male 's thick mane. The coloring material of the male 's mane varies from blond to black, by and large going darker as the king of beasts grows older. The most typical characteristic shared by both females and males is that the tail ends in a haired tussock. In some lions, the tussock conceals a difficult `` spinal column '' or `` spur '' , about 5 millimeters long, formed of the concluding subdivisions of tail bone fused together. The king of beasts is the lone feline to hold a caespitose tail – the map of the tussock and spinal column are unknown. Absent at birth, the tussock develops around 5 1⁄2 months of age and is readily identifiable at 7 months.

Nowak indicates the typical weight scope of lions as 150 to 250 kilograms ( 331 to 551 pound ) for males and 120 to 182 kilograms ( 265 to 401 pound ) for females. The size of grownup lions varies across their scope, with those from the southern African populations in Zimbabwe, the Kalahari and Kruger Park ( P. l. bleyenberghi, P. l. vernayi, or P. l. krugeri ) averaging about 189.6 kilogram ( 418 pound ) and 126.9 kilogram ( 280 pound ) in males and females severally, compared to 174.9 kilograms ( 386 pound ) and 119.5 kilogram ( 263 pound ) of male and female lions from East Africa. Reported organic structure measurings in males are head-body lengths runing from 170 to 250 centimeter ( 5 ft 7 in to 8 foots 2 in ) , tail lengths of 90–105 centimeter ( 2 ft 11 in–3 foot 5 in ) . In females reported head-body lengths range from 140 to 175 centimeter ( 4 ft 7 in to 5 foots 9 in ) , tail lengths of 70–100 centimeter ( 2 ft 4 in–3 foot 3 in ) , nevertheless, the often cited maximal caput and organic structure length of 250 centimeter ( 8 ft 2 in ) fits instead to extinct Pleistocene signifiers, like the American king of beasts, with even big modern lions mensurating several centimeters less in length.


The mane of the grownup male king of beasts, alone among cats, is one of the most typical features of the species. In rare instances a female king of beasts can hold a mane. The presence, absence, coloring material, and size of the mane is associated with familial stipulation, sexual adulthood, clime, and testosterone production ; the regulation of pollex is the darker and fuller the mane, the healthier the king of beasts. Sexual choice of couples by lionesses favors males with the densest, darkest mane. Research in Tanzania besides suggests mane length signals contending success in male–male relationships. Darker-maned persons may hold longer generative lives and higher offspring endurance, although they suffer in the hottest months of the twelvemonth.

Scientists one time believed that the distinguishable position of some races could be justified by morphology, including the size of the mane. Morphology was used to place races such as the Barbary king of beasts and Cape king of beasts. Research has suggested, nevertheless, that environmental factors such as mean ambient temperature influence the coloring material and size of a king of beasts 's mane. The ice chest ambient temperature in European and North American menagerie, for illustration, may ensue in a heavier mane. Thus the mane is non an appropriate marker for placing races. The males of the Asiatic races, nevertheless, are characterised by sparser Maness than mean African lions.

White lions

The white king of beasts is non a distinguishable races, but a particular morph with a familial status, leucism, that causes paler color akin to that of the white tiger ; the status is similar to melanosis, which causes black jaguars. They are non albinos, holding normal pigmentation in the eyes and tegument. White Transvaal king of beasts ( Panthera leo krugeri ) persons on occasion have been encountered in and around Kruger National Park and the next Timbavati Private Game Reserve in eastern South Africa, but are more normally found in imprisonment, where breeders intentionally select them. The unusual pick coloring material of their coats is caused by a recessionary allelomorph. Reportedly, they have been bred in cantonments in South Africa for usage as trophies to be killed during transcribed Hunts.

Group administration

Lions are the most socially inclined of all wild felines, most of which remain rather lone in nature. The king of beasts is a predatory carnivore with two types of societal organisation. Some lions are occupants, populating in groups of related lionesses, their couples, and offspring. Such a group is called a pride. Females form the stable societal unit in a pride and do non digest outside females. Membership lone alterations with the births and deceases of lionesses, although some females do go forth and go mobile. Although highly big prides, dwelling of up over 30 persons, have been observed, the mean pride consists may dwell of around 15 lions ; dwelling of several grownup females, up to four males ( known as a alliance if more than one ) and their greenhorn of both sexes. The exclusive exclusion to this form is the Tsavo king of beasts pride which ever has merely one grownup male. Male greenhorns are excluded from their maternal pride when they reach adulthood at around 2–3 old ages of age. The 2nd organisational behavior is labeled nomads, who range widely and travel about periodically, either singularly or in braces. Pairs are more frequent among related males who have been excluded from their birth pride. Note that a king of beasts may exchange life styles ; nomads may go occupants and frailty versa. Interactions between prides and nomads tend to be hostile, although pride females in estrous may let nomad males to near them.

The country a pride occupies is called a pride country, whereas that by a nomad is a scope. The males associated with a pride tend to remain on the peripheries, policing their district. Why sociality – the most marked in any cat species – has developed in lionesses is the topic of much argument. Increased runing success appears an obvious ground, but this is less than certain upon scrutiny: coordinated hunting does let for more successful predation but besides ensures that non-hunting members cut down per capita calorific consumption ; nevertheless, some take a function raising greenhorns, who may be left entirely for extended periods of clip. Members of the pride on a regular basis tend to play the same function in Hunts and hone their accomplishments. The wellness of the huntsmans is the primary demand for the endurance of the pride, and they are the first to devour the quarry at the site it is taken. Other benefits include possible blood-related choice ( better to portion nutrient with a related king of beasts than with a alien ) , protection of the immature, care of district, and single insurance against hurt and hungriness.

Both males and females can support the pride against interlopers, but the male king of beasts is better-suited for this intent due to its stockier, more powerful physique. Some persons systematically lead the defense mechanism against interlopers, while others lag behind. Lions tend to presume specific functions in the pride. Those dawdling buttocks may supply other valuable services to the group. An alternate hypothesis is that there is some wages associated with being a leader who fends off interlopers, and the rank of lionesses in the pride is reflected in these responses. The male or males associated with the pride must support their relationship to the pride from outside males who attempt to take over their relationship with the pride.

Hunting and diet

Lions are about wholly carnivorous and normally Hunt in groups. Their quarry consists chiefly of moderate-sized mammals, with a penchant for wildebeest, zebras, American bison, and warthogs in Africa and nilgai, wild Sus scrofa, and several cervid species in India. Many other species are hunted, based on handiness, chiefly ungulates weighing between 50 and 300 kilogram ( 110 and 660 pound ) such as koodoo, hartebeest, gemsbuck, and eland. They make besides take comparatively little species such as Thomson 's gazelle. Single lions are capable of conveying down quarries like zebra and wildebeest, which can be twice their ain weight, while runing larger quarry like camelopard and buffalo entirely are excessively much of a hazard. Cooperative-hunting lions are normally successful.

Extensive surveies show that lionesses usually prey on mammals with an mean weight of 126 kilograms ( 278 pound ) , while putting to deaths made by male lions average 399 kilogram ( 880 pound ) . In Africa, wildebeest rank at the top of preferable quarry ( doing about half of the king of beasts quarry in the Serengeti ) followed by zebra. Lions by and large avoid to the full adult grownup elephants, river horses, rhinoceroses, every bit good as really little quarries like dik-dik, coney, hare and monkey. However, camelopard and American bisons are frequently taken in certain parts. For case, in Kruger National Park, camelopard are on a regular basis hunted. In Manyara Park, Cape buffaloes represent every bit much as 62 % of the king of beasts 's diet, due to the high figure denseness of American bisons. Occasionally river horse is besides taken, but grownup rhinoceroses are by and large avoided. Warthogs are frequently taken depending on handiness. The lions of Savuti, Botswana, have adapted to runing immature elephants during the dry season, and a pride of 30 lions has been recorded killing persons between the ages of four and eleven old ages. In the Kalahari desert in South Africa, black-maned lions may trail baboons up a tree, delay patiently, so assail them when they try to get away. Lions besides attack domestic farm animal and in India cowss contribute significantly to their diet. Lions will kill other marauders such as leopards, chetah, and hyaenas, but they rarely devour them.

Young lions first show stalking behavior around three months of age, although they do non take part in runing until they are about a twelvemonth old. They begin to run efficaciously when approaching the age of two. In prides, lionesses do most of the hunting. In typical Hunts, each lioness has a favoured place in the group, either stalking quarry on the `` wing '' so assailing, or traveling a smaller distance in the Centre of the group and capturing quarry in flight from other lionesses. Cooperation is particularly of import when repressing and killing big quarries like American bison and camelopard. Males attached to prides do non normally take part in group hunting. However, some grounds suggests that pride males are merely every bit successful as females ; they are solo huntsmans who ambush prey in little shrub. Lions they are non peculiarly known for their staying power – for case, a lioness ' bosom makes up merely 0.57 % of her organic structure weight ( a male 's is about 0.45 % of his organic structure weight ) , whereas a hyaena 's bosom is near to 1 % of its organic structure weight. Therefore, they merely run fast in short explosions, and need to be near to their quarry before get downing the onslaught. They take advantage of factors that cut down visibleness ; many putting to deaths take topographic point near some signifier of screen or at dark.

The onslaught is short and powerful ; they attempt to catch the victim with a fast haste and concluding spring. The quarry normally is killed by choking, which can do intellectual ischaemia or asphyxia ( which consequences in hypoxemic, or `` general '' , hypoxia ) . The quarry besides may be killed by the king of beasts enveloping the animate being 's oral cavity and anterior nariss in its jaws ( which would besides ensue in asphyxia ) . Prey is typically eaten at the location of the Hunt, although big quarry is sometimes dragged into screen. Lions tend to quibble over a putting to death, peculiarly the males. When nutrient is scarce, cubs tend to endure the most but otherwise all pride members can eat their fill, including old and crippled 1s which can populate on leftovers. There is more sharing of larger putting to deaths. An grownup lioness requires an norm of about 5 kilograms ( 11 pound ) of meat per twenty-four hours, a male about 7 kilograms ( 15 pound ) . A king of beasts may ingurgitate itself and eat up to 30 kilograms ( 66 pound ) in one posing ; if it is unable to devour all the putting to death it will rest for a few hours before devouring more. On a hot twenty-four hours, the pride may withdraw to shadow go forthing a male or two to stand guard. Lions will support their putting to deaths from scavengers like vultures and hyaenas.

Lions and spotted hyaenas occupy the same ecological niche, intending they compete for quarry and carrion in the countries where they coexist. A reappraisal of informations across several surveies indicates a dietetic convergence of 58.6 % . Lions typically ignore spotted hyaena unless the lions are on a putting to death or are being harassed by the hyaenas, while the latter tend to visibly respond to the presence of lions whether there is nutrient or non. Lions seize the putting to deaths of patched hyaenas: in the Ngorongoro crater, it is common for lions to exist mostly on putting to deaths stolen from hyaenas, doing the hyaena to increase their kill rate. On the other manus, in Northern Botswana 's Chobe National Park, the state of affairs is reversed: hyaenas often challenge lions and steal their putting to deaths: they obtain nutrient from 63 % of all king of beasts putting to deaths. When confronted on a putting to death by lions, spotted hyaena may either go forth or wait patiently at a distance of 30–100 m ( 98–328 foot ) until the lions have finished, but they are besides bold plenty to feed alongside lions, and even coerce the lions off a putting to death. The two species may assail one another even when there is no nutrient involved for no evident ground. Lion predation can account for up to 71 % of hyena deceases in Etosha. Spotted hyaenas have adapted by often thronging lions that enter their districts. Experiments on confined spotted hyaenas revealed that specimens with no anterior experience with lions act indifferently to the sight of them, but will respond fearfully to the aroma. The size of male lions allows them on occasion to face hyaenas in otherwise equally matched bash and so to tip the balance in favor of the lions.

Lions tend to rule smaller felids such as chetahs and leopards where they co-occur, stealing their putting to deaths and killing their greenhorn and even grownups when given the opportunity. The chetah has a 50 % opportunity of losing its putting to death to lions or other marauders. Lions are major slayers of chetah greenhorns, accounting for up to 78.2 % of predator-killed juveniles in one survey. Cheetahs avoid competition by runing at different times of the twenty-four hours and conceal their greenhorns in thick coppice. Leopards besides use such tactics, but have the advantage of being able to exist much better on little quarry than either lions or chetah. Besides, unlike chetah, leopards can mount trees and utilize them to maintain their greenhorns and kills off from lions ; nevertheless, lionesses will on occasion be successful in mounting to recover leopard putting to deaths. Similarly, lions dominate African wild Canis familiariss, non merely taking their putting to deaths but besides feeding on immature and ( seldom ) grownup Canis familiariss. Population densenesss of wild Canis familiariss are low in countries where lions are more abundant. However, there are a few reported instances of old and wounded lions falling quarry to wild Canis familiariss.

While lions do non normally hunt people, some ( normally males ) seem to seek out human quarry ; one well-publicised instance includes the Tsavo maneaters, where 28 officially recorded railroad workers constructing the Kenya-Uganda Railway were taken by lions over nine months during the building of a span over the Tsavo River in Kenya in 1898. The huntsman who killed the lions wrote a book detailing the animate beings ' predatory behavior. The lions were larger than normal, lacked Maness, and one seemed to endure from tooth decay. The frailty theory, including tooth decay, is non favoured by all research workers ; an analysis of dentitions and jaws of man-eating lions in museum aggregations suggests that while tooth decay may explicate some incidents, prey depletion in human-dominated countries is a more likely cause of king of beasts predation on worlds.

The king of beasts 's propensity for man-eating has been consistently examined. American and Tanzanian scientists report that man-eating behavior in rural countries of Tanzania increased greatly from 1990 to 2005. At least 563 villagers were attacked and many eaten over this period – a figure far transcending the more celebrated `` Tsavo '' incidents of a century earlier. The incidents occurred near Selous National Park in Rufiji District and in Lindi Province near the Mozambican boundary line. While the enlargement of villagers into bush state is one concern, the writers argue that preservation policy must extenuate the danger because, in this instance, preservation contributes straight to human deceases. Cases in Lindi have been documented where lions seize worlds from the centre of significant small towns. Another survey of 1,000 people attacked by lions in southern Tanzania between 1988 and 2009 found that the hebdomads following the full Moon ( when there was less moonlight ) were a strong index of increased dark onslaughts on people.

A man-eating king of beasts was killed by game lookouts in Southern Tanzania in April 2004. It is believed to hold killed and eaten at least 35 people in a series of incidents covering several small towns in the Rufiji Delta coastal part. Dr Rolf D. Baldus, the GTZ wildlife programme coordinator, commented that it was likely that the king of beasts preyed on worlds because it had a big abscess underneath a grinder that was cracked in several topographic points. He farther commented that `` This king of beasts likely experienced a batch of hurting, peculiarly when it was masticating. '' GTZ is the German development cooperation bureau and has been working with the Tanzanian authorities on wildlife preservation for about two decennaries. As in other instances this king of beasts was big, lacked a mane, and had a tooth job.

Reproduction and life rhythm

The mean gestation period is around 110 yearss, the female giving birth to a litter of one to four greenhorns in a privy lair ( which may be a brush, a reed-bed, a cave, or some other sheltered country ) normally off from the remainder of the pride. She will frequently run by herself while the greenhorns are still incapacitated, remaining comparatively close to the brush or lair where the greenhorn are kept. The greenhorn themselves are born blind – their eyes do non open until approximately a hebdomad after birth. They weigh 1.2–2.1 kilogram ( 2.6–4.6 pound ) at birth and are about incapacitated, get downing to creep a twenty-four hours or two after birth and walking about three hebdomads of age. The lioness moves her greenhorn to a new den site several times a month, transporting them one by one by the scruff of the cervix, to forestall aroma from constructing up at a individual lair site and therefore avoiding the attending of marauders that may harm the greenhorn.

Normally, the female parent does non incorporate herself and her greenhorn back into the pride until the greenhorns are six to eight hebdomads old. Sometimes this debut to plume life occurs earlier, nevertheless, peculiarly if other lionesses have given birth at about the same clip. For case, lionesses in a pride frequently synchronise their generative rhythms and there is communal raising and suckling of the immature ( once the greenhorns are past the initial phase of isolation with their female parent ) , who suckle randomly from any or all of the nursing females in the pride. The synchronism of births besides has an advantage in that the greenhorns end up being approximately the same size, and therefore hold an equal opportunity of endurance, and older greenhorns do non rule the sucklings.

Weaning occurs after six to seven months. Male lions reach adulthood at about 3 old ages of age and, at 4–5 old ages of age, are capable of disputing and displacing the grownup male ( s ) associated with another pride. They begin to age and weaken between 10 and 15 old ages of age at the latest, Furthermore, when one or more new males oust the old male ( s ) associated with a pride, the vanquisher ( s ) frequently kill any bing immature greenhorns, possibly because females do non go fertile and receptive until their cubs mature or dice. A lioness frequently will try to support her greenhorns ferociously from a usurping male, but such actions are seldom successful. Success is more likely when a group of three or four female parents within a pride articulation forces against one male. Other beginnings of mortality for greenhorns include famishment and forsaking every bit good such as predation by leopards, hyaenas and wild Canis familiariss. All in all, every bit many as 80 % of the greenhorn will decease before the age of two.


Assorted species of tick normally infest the ears, cervix and inguen parts of most lions. Adult signifiers of several species of the cestode genus Taenia have been isolated from bowels, the lions holding ingested larval signifiers from antelope meat. Lions in the Ngorongoro Crater were afflicted by an eruption of stable fly ( Stomoxys calcitrans ) in 1962 ; this resulted in lions going covered in bloody bare spots and emaciated. Lions sought unsuccessfully to hedge the biting flies by mounting trees or creeping into hyena tunnels ; many perished or emigrated as the population dropped from 70 to 15 persons. A more recent eruption in 2001 killed six lions. Lions, particularly in imprisonment, are vulnerable to the eyetooth distemper virus ( CDV ) , feline immunodeficiency virus ( FIV ) , and feline infective peritoneal inflammation ( FIP ) . CDV is spread through domestic Canis familiariss and other carnivores ; a 1994 eruption in Serengeti National Park resulted in many lions developing neurological symptoms such as ictuss. During the eruption, several lions died from pneumonia and phrenitis. FIV, which is similar to HIV while non known to adversely affect lions, is unreassuring plenty in its consequence in domestic cats that the Species Survival Plan recommends systematic testing in confined lions. It occurs with high to endemic frequence in several wild king of beasts populations, but is largely absent from Asiatic and Namibian lions.


When resting, king of beasts socialization occurs through a figure of behaviors, and the animate being 's expressive motions are extremely developed. The most common peaceable haptic gestures are head rubbing and societal defeat, which have been compared with preparing in Primatess. Head rubbing – cuddling one 's brow, face and cervix against another king of beasts – appears to be a signifier of salutation, as it is seen frequently after an animate being has been apart from others, or after a battle or confrontation. Males tend to rub other males, while greenhorns and females rub females. Social creaming frequently occurs in tandem with caput friction ; it is by and large common and the receiver appears to show pleasance. The caput and cervix are the most common parts of the organic structure licked, which may hold arisen out of public-service corporation, as a king of beasts can non cream these countries separately.

Their repertory of voices is besides big ; fluctuations in strength and pitch, instead than distinct signals, appear cardinal to communicating. Most lion vocals are fluctuations of growling/snarling, miaouing and howling. Other sounds produced include whizzing, whiffing, bleating and humming. Lions tend to howl in a really characteristic mode, get downing with a few deep, long booms that trail off into a series of shorter 1s. They most frequently roar at dark ; the sound, which can be heard from a distance of 8 kilometers ( 5.0 myocardial infarction ) , is used to publicize the animate being 's presence. Lions have the loudest boom of any large cat.

Distribution and home ground

In Africa, lions can be found in savanna grasslands with scattered Acacia trees, which serve as shadiness ; their home ground in India is a mixture of dry savanna forest and really dry deciduous chaparral wood. The home ground of lions originally spanned the southern parts of Eurasia, runing from Greece to India, and most of Africa except the cardinal rainforest-zone and the Sahara desert. Herodotus reported that lions had been common in Greece in 480 BC ; they attacked the luggage camels of the Persian male monarch Xerxes on his March through the state. Aristotle considered them rare by 300 BC. By 100 AD they were extirpated. A population of Asian lions survived until the 10th century in the Caucasus, their last European outstation.

The species was eradicated from Palestine by the Middle Ages and from most of the remainder of Asia after the reaching of readily available pieces in the 18th century. Between the late nineteenth and early twentieth centuries, they became nonextant in North Africa and Southwest Asia. By the late 19th century, the king of beasts had disappeared from Turkey and most of northern India, while the last sighting of a unrecorded Asiatic king of beasts in Iran was in 1941 ( between Shiraz and Jahrom, Fars Province ) , although the cadaver of a lioness was found on the Bankss of the Karun river, Khūzestān Province in 1944. There are no subsequent dependable studies from Iran. The races now survives merely in and around the Gir Forest of northwesterly India. Approximately 500 lions live in the country of the 1,412 km2 ( 545 sq myocardial infarction ) sanctuary in the province of Gujarat, which covers most of the forest. Their Numberss have increased from 180 to 523 animate beings chiefly because the natural quarry species have recovered.

Population and preservation position

Most lions now live in eastern and southern Africa, and their Numberss there are quickly diminishing, with an estimated 30–50 % diminution per 20 old ages in the late half of the 20th century. Estimates of the African king of beasts population scope between 16,500 and 47,000 life in the natural state in 2002–2004, down from early 1990s estimations that ranged every bit high as 100,000 and possibly 400,000 in 1950. Primary causes of the diminution include disease and human intervention. Habitat loss and struggles with worlds are considered the most important menaces to the species. The staying populations are frequently geographically isolated from one another, which can take to inbreeding, and accordingly, reduced familial diverseness. Therefore, the king of beasts is considered a vulnerable species by the International Union for Conservation of Nature, while the Asiatic races is endangered. The king of beasts population in the part of West Africa is isolated from lion populations of Central Africa, with small or no exchange of engendering persons. The figure of mature persons in West Africa is estimated by two separate recent studies at 850–1,160 ( 2002/2004 ) . There is dissension over the size of the largest single population in West Africa: the estimations range from 100 to 400 lions in Burkina Faso 's Arly-Singou ecosystem. Another population in northwesterly Africa is found in Waza National Park, where about 14–21 animate beings persist.

Conservation of both African and Asiatic lions has required the apparatus and care of national Parkss and game militias ; among the best known are Etosha National Park in Namibia, Serengeti National Park in Tanzania, and Kruger National Park in eastern South Africa. The Ewaso Lions Project protects lions in the Samburu National Reserve, Buffalo Springs National Reserve and Shaba National Reserve of the Ewaso Ng'iro ecosystem in Northern Kenya. Outside these countries, the issues originating from lions ' interaction with farm animal and people normally consequences in the riddance of the lions. In India, the last safety of the Asiatic king of beasts is the 1,412 km2 ( 545 sq myocardial infarction ) Gir Forest National Park, and environing countries, in western India, which had about 180 lions in 1974 and about 400 in 2010. As in Africa, legion human habitations are close by with the end point jobs between lions, farm animal, locals and wildlife functionaries. The constitution of a 2nd independent population of Asian lions is planned at the Kuno Wildlife Sanctuary in the Indian province of Madhya Pradesh.

The former popularity of the Barbary king of beasts as a menagerie animate being has meant that scattered lions in imprisonment are likely to be descended from Barbary king of beasts stock. This includes lions at Port Lympne Wild Animal Park in Kent, England that are descended from animate beings owned by the King of Morocco. Another 11 animate beings believed to be Barbary lions were found in Addis Ababa menagerie, posterities of animate beings owned by Emperor Haile Selassie. WildLink International, in coaction with Oxford University, launched their ambitious International Barbary Lion Project with the purpose of identifying and genteelness Barbary lions in imprisonment for eventual reintroduction into a national park in the Atlas Mountains of Morocco.

Following the find of the diminution of king of beasts population in Africa, several coordinated attempts affecting king of beasts preservation have been organised in an effort to stem this diminution. Lions are one species included in the Species Survival Plan, a coordinated effort by the Association of Zoos and Aquariums to increase its opportunities of endurance. The program was originally started in 1982 for the Asiatic king of beasts, but was suspended when it was found that most Asian lions in North American menagerie were non genetically pure, holding been hybridised with African lions. The African king of beasts program started in 1993, concentrating particularly on the South African races, although there are troubles in measuring the familial diverseness of confined lions, since most persons are of unknown beginning, doing care of familial diverseness a job.

In imprisonment

Lions are portion of a group of alien animate beings that are the nucleus of menagerie exhibits since the late 18th century ; members of this group are constantly big craniates and include elephants, rhinoceroses, river horses, big Primatess, and other large cats ; zoos sought to garner as many of these species as possible. Although many modern menageries are more selective about their exhibits, there are more than 1,000 African and 100 Asian lions in menagerie and wildlife Parkss around the universe. They are considered an embassador species and are kept for touristry, instruction and preservation intents. Lions can make an age of over 20 old ages in imprisonment ; Apollo, a resident king of beasts of Honolulu Zoo in Honolulu, Hawaii, died at age 22 in August 2007. His two sisters, born in 1986, were still alive in August 2007. Engendering plans need to observe beginnings to avoid engendering different races and therefore cut downing preservation value. However, several Asiatic-African king of beasts crosses have been bred.

At the ancient Egyptian metropoliss of Taremu and Per-Bast were temples to the lioness goddesses of Egypt, Sekhmet and Bast and at Taremu there was a temple to the boy of the divinity, Maahes the king of beasts prince, where unrecorded lions were kept and allowed to roll within his temple. The Greeks called the metropolis Leontopolis, the `` City of Lions '' and documented that pattern. Lions were kept and bred by Assyrian male monarchs every bit early as 850 BC, and Alexander the Great was said to hold been presented with tame lions by the Malhi of northern India. Later in Roman times, lions were kept by emperors to take portion in the gladiator sphere or for executings ( see bestiarii, damnatio ad bestias, and venatio ) . Roman luminaries, including Sulla, Pompey, and Julius Caesar, frequently ordered the mass slaughter of 100s of lions at a clip. In the East, lions were tamed by Indian princes, and Marco Polo reported that Kublai Khan kept lions inside. The first European `` menagerie '' spread among baronial and royal households in the 13th century, and until the 17th century were called harems ; at that clip, they came to be called menageries, an extension of the cabinet of wonders. They spread from France and Italy during the Renaissance to the remainder of Europe. In England, although the seraglio tradition was less developed, lions were kept at the Tower of London in a harem established by King John in the 13th century, likely stocked with animate beings from an earlier menagerie started in 1125 by Henry I at his runing lodge in Woodstock, near Oxford ; where lions had been stocked harmonizing to William of Malmesbury.

Harems served as looks of the aristocracy 's power and wealth. Animals such as large cats and elephants, in peculiar, symbolised power, and would be pitted in battles against each other or domesticated animate beings. By extension, menageries and harems served as presentations of the laterality of humanity over nature. Consequently, the licking of such natural `` Godheads '' by a cow in 1682 astonished the witnesss, and the flight of an elephant before a rhinoceros drew mockeries. Such battles would easy melt out in the 17th century with the spread of the menagerie and their appropriation by the common mans. The tradition of maintaining large cats as pets would last into the 19th century, at which clip it was seen as extremely bizarre.

The presence of lions at the Tower of London was intermittent, being restocked when a sovereign or his consort, such as Margaret of Anjou the married woman of Henry VI, either sought or were given animate beings. Records indicate they were kept in hapless conditions at that place in the 17th century, in contrast to more unfastened conditions in Florence at the clip. The menagerie was unfastened to the populace by the 18th century ; admittance was a amount of three half-pence or the supply of a cat or Canis familiaris for feeding to the lions. A rival menagerie at the Exeter Exchange besides exhibited lions until the early 19th century. The Tower menagerie was closed down by William IV, and animate beings transferred to the London Zoo, which opened its Gatess to the populace on 27 April 1828.

The wild animate beings trade flourished aboard improved colonial trade of the 19th century. Lions were considered reasonably common and cheap. Although they would barter higher than Liberation Tigers of Tamil Eelams, they were less dearly-won than larger, or more hard to transport animate beings such as the camelopard and river horse, and much less than elephantine coon bear. Like other animate beings, lions were seen as little more than a natural, unbounded trade good that was pitilessly exploited with awful losingss in gaining control and transit. The widely reproduced imagination of the heroic huntsman trailing lions would rule a big portion of the century. Explorers and huntsmans exploited a popular Manichean division of animate beings into `` good '' and `` evil '' to add thrilling value to their escapades, projecting themselves as epic figures. This resulted in large cats, ever suspected of being cannibals, stand foring `` both the fright of nature and the satisfaction of holding overcome it. ''

Lions were kept in cramped and seamy conditions at London Zoo until a larger king of beasts house with roomier coops was built in the 1870s. Further alterations took topographic point in the early 20th century, when Carl Hagenbeck designed enclosures more closely resembling a natural home ground, with concrete 'rocks ' , more unfastened infinite and a fosse alternatively of bars. He designed king of beasts enclosures for both Melbourne Zoo and Sydney 's Taronga Zoo, among others, in the early 20th century. Though his designs were popular, the old bars and coop enclosures prevailed until the sixtiess in many menagerie. In the ulterior decennaries of the 20th century, larger, more natural enclosures and the usage of wire mesh or laminated glass alternatively of lowered lairs allowed visitants to come closer than of all time to the animate beings, with some attractive forces even puting the lair on land higher than visitants, such as the Cat Forest/Lion Overlook of Oklahoma City Zoological Park. Lions are now housed in much larger realistic countries ; modern recommended guidelines more closely approximative conditions in the wild with closer attending to the lions ' demands, foregrounding the demand for lairs in separate countries, elevated places in both Sun and shadiness where lions can sit and adequate land screen and drainage every bit good as sufficient infinite to roll. There have besides been cases where a king of beasts was kept by a private person, such as the lioness Elsa, who was raised by George Adamson and his married woman Joy Adamson and came to develop a strong bond with them, peculiarly the latter. The lioness subsequently achieved celebrity, her life being documented in a series of books and movies.

Baiting and chastening

Lion chastening refers to the pattern of chastening lions for amusement, either as portion of an established circus or as an single act, such as Siegfried & Roy. The term is besides frequently used for the taming and show of other large cats such as Liberation Tigers of Tamil Eelams, leopards, and pumas. The pattern was pioneered in the first half of the 19th century by Frenchman Henri Martin and American Isaac Van Amburgh who both toured widely, and whose techniques were copied by a figure of followings. Van Amburgh performed before Queen Victoria in 1838 when he toured Great Britain. Martin composed a dumb show titled Les Lions de Mysore ( `` the lions of Mysore '' ) , an thought that Amburgh rapidly borrowed. These Acts of the Apostless eclipsed equestrianism Acts of the Apostless as the cardinal show of circus shows, but genuinely entered public consciousness in the early 20th century with film. In showing the high quality of homo over animate being, king of beasts chastening served a purpose similar to carnal battles of old centuries. The ultimate cogent evidence of a tamer 's laterality and control over a king of beasts is demonstrated by puting his caput in the king of beasts 's oral cavity. The now iconic king of beasts tamer 's chair was perchance foremost used by American Clyde Beatty ( 1903–1965 ) .

Cultural significance

Representations of lions day of the month back to the early Upper Paleolithic. The lioness-headed tusk carving from Vogelherd cave in the Swabian Alb in southwesterly Germany, dubbed Löwenmensch ( lion-human ) in German. The sculpture has been determined to be at least 32,000 old ages old from the Aurignacian civilization, but it may day of the month to every bit early as 40,000 old ages ago. The sculpture has been interpreted as anthropomorphous, giving human features to an animate being, nevertheless, it besides may stand for a divinity. Two lions were depicted coupling in the Chamber of Felines in 15,000-year-old Paleolithic cave pictures in the Lascaux caves. Cave lions besides are depicted in the Chauvet Cave, discovered in 1994 ; this has been dated at 32,000 old ages of age, though it may be of similar or younger age to Lascaux.

In Africa, cultural positions of the king of beasts have varied by part. In some civilizations, the king of beasts symbolises power and royalty and some powerful swayers had the word `` king of beasts '' in their moniker. For illustration, Marijata of the Mali Empire ( c. 1235–c. 1600 ) was given the name `` Lion of Mali '' . Njaay, the legendary laminitis of the Waalo land ( 1287–1855 ) , is said to hold been raised by lions and returned to his people part-lion to unify them utilizing the cognition he learned from the animals. In much of West Africa, to be compared to a king of beasts was considered to be one of the greatest regards. The societal hierarchies of their societies where connected to the carnal land and the king of beasts represented the top category. In parts of West and East Africa, the king of beasts is associated with healing and is seen as the nexus between the visionaries and the supernatural. In other East African traditions, the king of beasts is the symbol of indolence. In many folk tales, lions are portrayed as holding low intelligence and are easy tricked by other animate beings.

In the Talmud, Chullin 59b, Rabbi Joshua ben Hananiah tells Emperor Hadrian about the elephantine king of beasts of the wood of Bei Ilai called the Tigris, a king of beasts so immense that the infinite between its ears measures 9 cubits. The emperor asks the rabbi to name Forth this king of beasts. He reluctantly agrees. At a distance of 400 parasangs from Rome it roars, and all pregnant adult females miscarry and all the walls of Rome fall down. Then it comes to 300 parasangs and booms, and all the forepart dentition and grinders of Roman work forces fall out, and even the emperor himself falls from his throne. He begs the rabbi to direct it back. The rabbi prays and it returns to its topographic point.

In the Puranic texts of Hinduism, Narasimha ( `` man-lion '' ) a half-lion, half-man embodiment or ( embodiment ) of Vishnu, is worshipped by his fans and saved the kid fan Prahlada from his male parent, the evil devil male monarch Hiranyakashipu ; Vishnu takes the signifier of half-man/half-lion, in Narasimha, holding a human trunk and lower organic structure, but with a lion-like face and claws. Singh is an ancient Indian vedic name intending `` king of beasts '' ( Asiatic king of beasts ) , dating back over 2000 old ages to ancient India. It was originally merely used by Rajputs a Hindu Kshatriya or military caste in India. After the birth of the Khalsa brotherhood in 1699, the Sikhs besides adopted the name `` Singh '' due to the wants of Guru Gobind Singh. Along with 1000000s of Hindu Rajputs today, it is besides used by over 20 million Sikhs worldwide. Found famously on legion flags and coats of weaponries all across Asia and Europe, the Asiatic lions besides stand house on the National Emblem of India. Farther South in South Asia, the Asiatic king of beasts is symbolic for the Sinhalese, Sri Lanka 's cultural bulk ; the term derived from the Indo-Aryan Sinhala, intending the `` king of beasts people '' or `` people with king of beasts blood '' , while a sword-wielding king of beasts is the cardinal figure on the national flag of Sri Lanka.

The Asiatic king of beasts is a common motive in Chinese art. They were foremost used in art during the late Spring and Autumn period ( 5th or 6th century BC ) , and became much more popular during the Han Dynasty ( 206 BC – AD 220 ) , when imperial guardian lions started to be placed in forepart of imperial castles for protection. Because lions have ne'er been native to China, early word pictures were slightly unrealistic ; after the debut of Buddhist art to China in the Tang Dynasty ( after the 6th century AD ) , lions normally were wingless, with shorter, thicker organic structures, and curly Maness. The king of beasts dance is a signifier of traditional dance in Chinese civilization in which performing artists mimic a king of beasts 's motions in a king of beasts costume, frequently with musical concomitant from cymbals, membranophones, and tam-tams. They are performed at Chinese New Year, the August Moon Festival and other celebratory occasions for good fortune.

`` Lion '' was the moniker of several mediaeval warrior swayers with a repute for courage, such as the English King Richard the Lionheart, Henry the Lion, ( German: Heinrich der Löwe ) , Duke of Saxony, William the Lion, King of Scotland, and Robert III of Flanders nicknamed `` The Lion of Flanders '' —a major Flemish national icon up to the present. Lions are often depicted on coats of weaponries, either as a device on shields themselves, or as protagonists, but the lioness is much more infrequent. The formal linguistic communication of heraldry, called coat of arms, employs Gallic footings to depict the images exactly. Such descriptions specified whether lions or other animals were `` rampant '' or `` passant '' , that is whether they were rise uping or stooping. The king of beasts is used as a symbol of featuring squads, from national association football squads such as England, Scotland and Singapore to famous nines such as the Detroit Lions of the NFL, Chelsea and Aston Villa of the English Premier League, ( and the Premiership itself ) , Eintracht Braunschweig of the Bundesliga, and to a host of smaller nine around the universe.

Lions continue to be featured in modern literature, from the messianic Aslan in The Lion, the Witch and the Wardrobe and following books from The Chronicles of Narnia series written by C. S. Lewis, to the comedic Cowardly Lion in The Wonderful Wizard of Oz. The coming of traveling images saw the continued presence of king of beasts symbolism ; one of the most iconic and widely recognized lions is Leo the Lion, which has been the mascot for Metro-Goldwyn-Mayer ( MGM ) studios since the 1920s. The 1960s saw the visual aspect of what is perchance the most celebrated lioness, the Kenyan animate being Elsa in the film Born Free, based on the true-life book of the same rubric. The king of beasts 's function as male monarch of the animals has been used in sketchs, such as the 1994 Disney animated characteristic movie The Lion King.

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